What The All-Father Means


Readers of this weblog may sometimes notice that I break out in pompous and self-important declarations of being a “scion of the All-Father.” This is basically a joke. But, it’s a joke that draws from a legitimate basis of science and mythology. The “All-Father” is another name for Odin. I’m really talking about Indra, who is probably more like Thor. And obviously, Norse paganism is only distantly related to the mythology of the Indo-Aryans. As someone more familiar with the lineaments of Northern European mythology than Indian, of course, it’s easier for me to draw on the motifs of the former to relate to the latter.

R1a distribution

The scientific component has to do with R1a. Specifically, R1a1a, defined by the M17 mutation (discovered by my boss at my day-job 20 years ago). There are two very closely related “clades,” that is, families of pedigrees, of this Y chromosomal lineage, passed from father to son. One of them defines mostly European R1a1a, Eastern Europeans, and to a lesser extent Western Europeans. Another branch is found mostly in Central and South Asia.

When I first saw this distribution around the year 2000 it left me scratching my head. Of course, I knew about the Indo-European languages. But I had always assumed that the demographic impact of the original Indo-Europeans was relatively marginal. And yet this Y chromosome was found at frequencies in the 10-50% range across vast swaths of Eurasia.

Much of the 2000s was spent on arguments as to whether R1a was indigenous to South Asia or to Central Eurasia. Ultimately these arguments were not resolvable due to limitations of the data. To calibrate dates and diversity researchers relied on microsatellites, which are useful due to their high mutation rates, but also erratic for the same reason (not only were confidence intervals wide, some of the assumptions of the model parameters were guesses).

In the early 2010s, whole-genome sequences of Y chromosomes came online. It became very clear that the most common R1a1a lineages exhibited the “star phylogeny.” Demographically, what this means is that men carrying this lineage underwent very rapid population expansion for a short period of time. So rapid that a “father” lineage would give rise to numerous “son” lineages one mutational step away

You can see in the figure that node “A” has given rise to a “star phylogeny.” A large number of individuals are one mutational step away from that genotype. A more normal phylogeny would produce a complex structured tree which accrues mutations across the various branches gradually.

In the South Asian context, a paper from 2004, Independent origins of Indian caste and tribal paternal lineages, introduced a result which prefigured what we now know:

Analyses of molecular variance also suggest that caste groups are more homogeneous for Y chromosome variation than tribal groups, since the variance among caste groups (sampled from all over India) is 3-fold less than that observed among tribal groups and 2-fold less than that observed among all Indian populations grouped together (Table 3). Moreover, if only north caste groups are considered, the variance among populations is not significantly different from zero (Table 3), indicating that spread over the Indian subcontinent although they are located up to ∼1500 km away from each other, these populations have highly homogeneous Y chromosome compositions.

The implications of the lack of structure of R1a on the Indo-Gangetic plain is always something that struck me. It suggested that the paternal lineages only recently expanded since they didn’ have time to build up distinct regional mutations. In contrast, the adivasi populations had a wider distribution of Y chromosomal haplogroups, and they exhibit a lot deeper diverged lineages.

Which brings me to the personal angle. In the spring of 2010, I did my first personal genomic test. I got my Y and mtDNA results back first. It turned out my Y was R1a1a, and my mtDNA was U2b. I was surprised by both. Eastern Bengali has the highest fraction of mtDNA macrohaplogroup M in the world. R1a1a was less surprising. But, it was very strange to have a concrete, personal, connection to this lineage which had been on my mind for a decade or so.

My funny attachment to my haplogroup is probably a function of my upbringing. Growing up as brown in the United States, I wasn’t exposed to Indian culture, nor was I well versed in the details of South Asian communalism. My family is pretty conventional in being upper-middle-class Bengali Muslims, so there is not a jati identity or anything like that I could identify with (and though my parents are Muslim, they are not extremely so, therefore religious identity was a background and not foreground variable). When I looked at my overall genome in 2010 it was clear I didn’t have the “runs of homozygosity” that characterize many people from South Asian backgrounds who come from endogamous communities. I know some of my ancestors were Kayasthas, and my father has some Brahmin ancestry, but the most distinctive thing about me in hindsight is I’m a typical east Bengali with more than a usual dollop of East Asian ancestry (my family is from Comilla).

My Y chromosomal haplogroup, in contrast, is something clear, distinct, and precise. It is an anchor, something which I use to channel my preoccupations and concerns. I don’t have Omar’s Gujar tribal ancestry, or Zach’s Muhajir/Persian origins. I’m just a brown American whose parents did not instill him a patriotism about the “motherland” (Bangladesh), because they themselves didn’t even live a decade in that nation. Though there is a spectrum, it is clear that many South Asian Americans are less “coconut” than I am, and are attuned to fine differences of status, origin, and background. Growing up around only white people my identity was racialized, not ethnicized.

I have never felt superior or inferior to any community or ethnicity of South Asian because I never belonged to any community, have weak ethnic identity, and don’t believe in any religion. The religious prejudices I do have are probably Anglo-Protestant ones against Catholicism, because of the implicit assumptions and background facts of America’s Whig culture.

What R1a1a symbolizes to me is that I have a concrete connection to a semi-historical phenomenon between the end of prehistory and before the written word, which we have not grasped or understood very well. Though it is true R1a1a is found at higher concentrations in “upper castes,” as well as in the north and west of the subcontinent, and among Indo-Aryan speakers, the reality is it is found in almost every community in South Asia (the main exception being among Tibeto-Burmans and Munda). There are many communities, such as Chenchus, which have very little steppe ancestry but retain a substantial proportion of R1a1a.

For obvious reasons this haplogroup is associated with Indo-Aryans (the earliest find of R1a1a-Z93 is from the Bronze Age Volga Srubna culture), but its reach is far beyond current areas of Indo-Aryan speech. Its ubiquity is a testament to a broader South Asia cultural matrix that emerged in the centuries after 1500 BC, from north to south.

This is of course not a moral judgment. The expansion of this paternal lineage at the expense of others likely occurred through a process of aggression and social exclusion. This is nothing to be proud of…or ashamed of. It’s just a description.

4+

Genetic variation across many South Asian communities

Someone in the comments posted the results from The Genomic Formation of South and Central Asia. I put the percentages with a few ratios in a Google doc. I don’t know what a lot of these groups are. Can readers illuminate? We need to be careful about the sample size, but I think there are a lot of interesting patterns in there.

Remember that “Steppe”, “Indus Periphery” and “Onge” are populations artifacts within a model. The way I explain it to people is that rather than focusing on the percentage, look at how the populations vary across the parameters. That is a pretty robust result. No matter what outgroups you’re going to use, Brahmins in most of South Asia seem to have more “West Eurasian” type ancestry than other populations (except in the NW). Because “Indus Periphery” has a minority of “Ancient Ancestral South Indians” (AASI) as part of its ancestry, the “Onge” fraction should be seen as a floor on AASI ancestry (the Onge ancestors diverged from the AASI ~40,000 years ago, so it’s a very large difference).

Continue reading “Genetic variation across many South Asian communities”

0

Lord Indra was a tan man

An angel of the Christian Era

I get a fair amount of email related to questions about Indian genetics, as well as calls for me to adjudicate various controversies. A major problem with any “Aryan invasion theory” or its descendants, which posit non-trivial gene flow from the Eurasian steppe, is the possibility that the Indo-Aryan ancestors of nearly all South Asians (albeit, in extremely varied proportions) were a thousand men with the bright faces, azure eyes, and flaxen locks. Paul Bettany times a thousand astride chariots.

The anachronistic neocolonialism obviously makes Indians uncomfortable. Or that’s my psychoanalysis. I don’t care much either way. What does Lord Indra’s scion care? We are the unbroken lineage, grasping Eurasia’s heart, from the Baltic to the Bay of Bengal!

The flip side of this is people of European ancestry, some of whom are white nationalists, do come close to making this claim. That is, that the Aryans were white Nordic people. The genetics from the Sintashta and Andronovo cultural complexes do indicate that they resemble many of the contemporaneous European populations. Their ultimate locus of origin probably is the Pontic steppe, which is in the geographic boundaries of Europe, as such. Finally, these steppe peoples exhibit genetic signatures of reflux from Europe. That is, they’re not just Yamna-descendants but derived from Yamna-like people who moved west, mixed with local indigenous Europeans, and moved back east along the Eurasian steppe corridor.

Lord Indra’s face? NO!

Looking at some of the ancient forensic DNA some of these individuals have suggested that I must admit that the Indo-Aryans were genetically like Europeans, and phenotypically like Europeans as well. They know that I won’t lie like some people, and just want me to admit this.

Continue reading “Lord Indra was a tan man”

0

The Syeds of South Asia are the sons of Hindus and Magians

The above figure shows the frequencies of Y chromosomal haplogroups of men of South Asian who claim to be descended from the prophet or his tribe, as cross-referend with their surnames. The “Non-IHL” category indicates those who are not of these honored lineages.

The paper from which I drew the data, Y chromosomes of self-identified Syeds from the Indian subcontinent show evidence of elevated Arab ancestry but not of a recent common patrilineal origin, actually somewhat support the idea that these people descend from Muhammad or the Quraysh or the Ansar.

I think this is wrong.

But first, why do think these data results show Arab affinity? The “IHL” lineages have a higher proportion of haplogroup J, the most common haplogroup among Arabs. J is not exactly rare in South Asia (lots of <<<Brahmins>>> who are not sons of Indra have it because they are the scions of cunning Dasa priests), but there’s clearly a frequency discrepancy.

And yet this paper was published in 2010. We now know through various tests of confirmed descendants of Muhammad, and who descend in the male line from his cousin Ali, that they carry a branch of haplogroup J1.

Even among the Syeds, most do not descend from Muhammad assuredly. There are nearly as many scions of Lord Indra, R1a1, as those who bear haplogroup J. Of the J’s within the Syed community, I think the most likely scenario if they are not South Asia is that they are Iranian. J is found at frequencies of 35% in Iran, and Iranians, along with Turks, were the most common migrants into South Asia.

In other words, the Syeds of the Indian subcontinent are the sons of magians, not Muhammad.

0

We are all Aryans now

Last year I contributed a chapter to a book soon to be published in India, Which of Us are Aryans? In answer to the question, the straightforward answer is that almost all of us are Aryans. That is, the thin but persistent layer of Indo-Aryan (“steppe”) ancestry is present across the subcontinent. In higher fractions among Brahmins and Kshatriyas than in Dalits, in the northwest than the southeast, and among Indo-European speakers than Dravidians. But this ancestral component and its cultural correlates are found across southern Asia.

Secondarily, there has been some discussion about the negative valence in the West about the term “Aryans.” In particular, its “cultural appropriation” by German Nazis by way of Theosophy and various spiritual and quasi-spiritual movements in the early 20th century.

As an American to see the word “Aryan” bandied about like this is strange and a bit uncomfortable. But there are now more than 1 billion Indians, so I don’t believe we in the West are a position dictate in terms of the lexicon that we borrowed from Indians in the first place, often without clear attribution (most Americans and Europeans would be surprised that “Aryan” is an Indian and Iranian term).

3+

The religious and genetic structure of Bengal & Partition

I was emailing with a friend of mine about population genetic history and Southeast Asia. I mentioned offhand that there is an east to west cline of Tibeto-Burman ancestry in Bengal. He expressed surprise, assuming Partition had scrambled everything.

As most readers of this weblog know, Partition was less traumatic for Bengal than it was for Punjab. The violence was less extreme, and the population movement also not as massive. And yet looking at the religious map it is clear that some sorting has occurred. The proportion of Hindus in the region that is now Bangladesh has gone from ~25% to about 10% over the past 70 years, or three generations. Though some of this is due to differences in fertility, the main driver has been migration of Hindus out of East Pakistan, and later Bangladesh. In contrast, there has not been much of a reciprocal migration of Muslims into Bangladesh.

This results in a peculiarity when I receive genotypes from people of Bengali origin: a large minority of people of Hindu background mention that one or both of their parents have origins in eastern Bengal, what is not Bangladesh. In contrast, I have never received a gentoype from someone who tells me that their family migrated from western Bengal into Bangladesh.

The genetic consequence is simple: there is a larger variance of East Asian ancestry in West Bengal than East Bengal because of more mixing in the west than the east. In contrast, one could probably infer the extent of the migration simply by doing genetic analysis and not looking at Census data!

0

The hammer of the All-Father


Unless you have been sleeping under a rock, a mildly slanderous piece in The New York Times Magazine has taken aim at David Reich and his band of paleogeneticists, Is Ancient DNA Research Revealing New Truths — or Falling Into Old Traps? I address this piece at my other weblog.

One of the major themes of the piece are the legends and myths of the people of Vanuatu:

I asked him about how the concept of Lapita migration to empty islands had been received by people whose oral traditions said they came from a stone or a coconut tree.

The reason this is relevant is that paleogeneticists have probed the history of Vanuatu. And yet this is the past. The future is that the Reich lab is collaborating with other paleogeneticists to crack the nut of the history of the Indian subcontinent with ancient DNA. They’ve been working on this for years, and they are working on it now. There are 275,000 people who live in Vanuatu. There are 1.7 billion people who live in the Indian subcontinent.

Within the next year I believe that the Reich lab will publish results which will falsify the beliefs of a substantial number of Indians about the nature of the origins of the native peoples of the region. This will shatter world-views, undermine mythologies, and rock peoples’ worlds. There will be sophists who live in denial, but the truth will be plain to those who see.

I understand that some of you reading this disagree with this assessment. Ultimately I don’t care because the data are coming, and if I’m wrong, that’s OK too. I don’t have emotional baggage invested in alternative models. But, I do wonder why the mythological traditions of “non-indigenous” people seem to warrant less attention than smaller nations or premodern tribes.

3+

How to avoid offending people?

Please watch the last three minutes of:

How to avoid very unexpectedly offending people when we don’t want to? How to have dialogue with people, ask them questions and get feedback from others without suddenly massively angering them?

This has nothing to do with Saira Roa’s actual opinions or high resolution fully integrated philosophy of philosophies. She seems to be a sweet loving person. Her perspective is unique and I would have loved to better understand it.

I have met many people from childhood who are suddenly and very unexpectedly massively triggered and angered. Often they will start accusing others of nazism, fascism, racism, bigotry, prejudice, sectarianism or some other related charge. In many cases immediately walk away. Many junior high school, high school, undergraduate and graduate level teachers at institutions I attended were this way. Some students were also this way, but truth be told teachers were far more likely to exhibit these symptoms than students. And a lot of the time, I and many others didn’t understand why this happened. Saira Roa is very middle of the road representative of very large numbers of people I have met (teachers and non teacher adults), (in the west or in India) and I am not picking on her. Rather I am asking how to avoid causing a massive firestorm when we don’t want to create one. In this case, Sargon didn’t want to anger her, but rather was very curious to better understand what she believes and why she believes what she believes.

This particular unexpected firestorm was set off when Sargon says to Saira Roa that some blacks were complicit in the slavery of other blacks. My questions about this is two fold:

  • Is there some way Sargon could have made a similar point without massively angering Saira Roa and causing her to end the interview?
  • Why did this statement elicit this reaction in the first place?

Saira Roa has a Hindu name. When the east (and large parts of Europe for that matter) was (were) conquered by Islamists (note that most muslims are not Islamists and today’s muslims are in no way responsible for the actions of their great ancestors), almost all eastern universities, libraries, temples, spiritual centers, scientific institutions etc. were destroyed. Much of the non muslim population was converted into slaves. Because of this, many Asian nonmuslims get emotional when the subject of slavery is mentioned. Could this be where part of Saira Roa’s feelings come from?

Most Asians (Indians included) and Africans initially welcomed Europeans as a way to drive Islamists out. Europeans as a quid pro quo of sorts banned slavery across Asia and Africa. This was deeply popular among nonmuslims and seen as sectarian Islamaphobia by many Islamists. [Obviously after this initial period, Africans and Asians wanted European colonizers to let them to be independent.] Perhaps Saira Rao thinks that the people who owned slaves on the African continent and sold them to South America, Central America, Mexico, Caribbean, North America, North Africa, East Africa, Europe, Asia were not really Africans but Islamist occupiers? Perhaps her definition of “African” or “black” is only nonmuslims with substantial sub-saharan African DNA haploid admixture? Therefore, “blacks” by her definition were not complicit in the slavery of other blacks and the exporting of black slaves around the world? I am not saying this is true. But rather could this be what she believes?

[Obviously some historians might posit the hypothesis that even if the large majority or vast majority of people who owned African slaves were muslim, at least some African slaves were owned by nonmuslims with substantial sub-saharan African DNA haploid admixture too. But perhaps Saira Roa disagrees with this.]

Are there other possible reasons for why she was so offended?

Can everyone reading please explain this to me in the comment section below? What advise does everyone have for how to avoid deeply angering or offending people in general? Thanks to everyone in advance.

2+

Are Haryana Jats the closest living descendents of our Vedic forefathers ?

Recently, there was a paper on some communities of Northwestern India such as Rors, Jats, Kambojs, Gujjars & Khatris. The primary focus of the paper was the community of cattle herders from Haryana known as Rors.

This is part 1 of my review of the paper. In part 2 I shall focus on whether the evidence furnished in the paper proves a steppe migration into South Asia.

Let me first quote the abstract in full :-

The Indus Valley has been the backdrop for several historic and prehistoric population movements between South Asia and West Eurasia. However, the genetic structure of present-day populations from Northwest India is poorly characterized. Here we report new genomewide genotype data for 45 modern individuals from four Northwest Indian populations, including the Ror, whose long-term occupation of the region can be traced back to the early Vedic scriptures. Our results suggest that although the genetic architecture of most Northwest Indian populations fits well on the broader North-South Indian genetic cline, culturally distinct groups such as the Ror stand out by being genetically more akin to populations living west of India; such populations include prehistorical and early historical ancient individuals from the Swat Valley near the Indus Valley. We argue that this affinity is more likely a result of genetic continuity since the Bronze Age migrations from the Steppe Belt than a result of recent admixture. The observed patterns of genetic relationships both with modern and ancient West Eurasians suggest that the Ror can be used as a proxy for a population descended from the Ancestral North Indian (ANI) population. Collectively, our results show that the Indus Valley populations are characterized by considerable genetic heterogeneity that has persisted over thousands of years.

Pay attention to the bolded part. As per the pre-print by Narasimhan et al, the ANI is the likely population that spread Steppe ancestry and hence Indo-Aryan ancestry among South Asians by mixing with  the ASI group. Now this paper on Rors says that Rors (by corollary the Jats) are the population most identical to this hypothetical ANI population. Please note – It is not Brahmins but a herder group from Haryana, which is the vert heartland of Vedic India. This is very significant because it clearly establishes the veracity of our Vedic tradition.

Let us look at this in more detail.

The ancestors of Rors and Jats from Haryana spread the Vedic civilization

As many of you here might be aware, the Vedic homeland was situated on the banks of the river Saraswati in a region which encompassed today’s Haryana and Western UP from where it eventually spread further into Northern India, principally in the Gangetic plains and beyond.

In terms of genetics therefore, one may argue that if there is a genetic signature of the Vedic people, it should be found most strongly in the original Vedic homeland and gradually reduce as one moves away from this homeland. Ofcourse, the caveat would be, that unless the modern people residing in the Vedic homeland had come to completely replace the original inhabitants of Haryana  who spread the Vedic culture.

The ancient DNA research has now shown that in terms of autosomal ancestry, there is link between the modern presence of Indo-European speakers across Eurasia and the ‘steppe’ ancestry component.

In South Asia it is argued, that the ‘steppe’ component is highest among the Brahmins and decreases as one moves down the caste heirarchy and this is said to be one of the principal evidences of movement of steppe people into South Asia having spread the Indo-European language and culture. Infact, the recent Narasimhan et al paper, even went so far as to suggest,

Although the enrichment for Steppe ancestry is not found in the southern Indian groups, the Steppe enrichment in the northern groups is striking as Brahmins and Bhumihars are among the traditional custodians of texts written in early Sanskrit. A possible explanation is that the influx of Steppe_MLBA ancestry into South Asia in the mid-2nd millennium BCE created a meta-population of groups with different proportions of Steppe ancestry, with ones having relatively more Steppe ancestry having a central role in spreading early Vedic culture.

However, it has already been known since many years that the population having the highest ‘steppe’ ancestry in South Asia are not the Brahmins but the Jats, more specifically, the Haryanvi Jats. This was also noted by Razib in one of his earlier blogs.

The present study focuses on this elevated steppe related component in Jats and more specifically in a related group from Haryana known as the Rors. It is titled, ” The Genetic Ancestry of Modern Indus Valley Populations from Northwest India “.  This study has the advantage that it incorporates the aDNA data from the Narasimhan et al and other recent papers.

The following is the admixture graph from the study,

As can be seen in the selected enlarged portion of the graph, the ‘steppe’ like light blue component, which is highest in some of the Northern European groups closest to the steppe, like the Latvians, Lithuanians, Russians etc., is far higher in Rors than it is in the Brahmins or any other South Asian group.

As per the authors themselves,

Outgroup f3 analysis in the form of (PNWI, X; Yoruba) showed that the Ror (and Jat) have distinct, high genetic similarity to modern Europeans (Figures 1C, 1D, and S5), far higher than the similarity observed in other NWI populations, such as the Gujjar (Figures 1D and S5). Among an extended set of South Asians, this pattern was repeated only in the Pathan population from Pakistan (Figure S5).

And,

Refined IBD analysis highlights the general trend whereby the sharing of IBD segments declines as one moves along the cline from PNWI and NI_IE toward Dravidian and Indian Austroasiatic (IN_AA) groups (Figure 2A). Strikingly, among all PNWI groups studied, the Ror demonstrate the highest number of IBD segments shared with Europeans and Central Asians, whereas the Gujjar share a higher number of IBD segments with local Indian Indo-Europeans and Dravidians than do other PNWI groups (Figure 2A).

In CHROMOPAINTER analysis, as expected, the Ror (and Jat) exhibited a significantly higher number of chunks received from Europeans than do other NWI populations studied (t test, p value < 0.01).

They also state further,

A higher level of European ancestry in the Ror and Jat compared to other South Asians (Figures 1, 2, S2, S5, and S13 and Tables S5–S8) makes these two populations outliers within the broader Northwest South Asian landscape. This could be indicative of either a possible recent gene flow from a population related to Europe or to ancient West-Eurasian-related influx, which would agree with previous studies on adaptation, wherein the Ror and Jat have stood out for their high frequency of the lactase persistence allele (LCT-13910T) and the light-skin-color gene variant (SLC24A5).

The Rors and Jats also have higher frequencies of Lactase persistence and light skin color gene variant which makes the case of their more recent ancestry sharing, compared to other South Asians, with Northern Europeans or steppe groups stronger.

Also,

We also report that, relative to other South Asians, the Ror group has high shared drift with the EHG and Steppe_EMBA groups, higher allele sharing with the Steppe_MLBA group, and higher affinity with the Iron Age (prehistorical) and early historical first South Asian ancient sources (Figures S6A, S6B, S7, S8A, S8D, and S9 and Tables S9 and S16).

Finally the authors argue that the Rors are the best proxy for the ANI ancestry in South Asians,

In summary, we demonstrate a higher proportion of genomic sharing between PNWI populations and ancient EHG and Steppe-related populations than we observe in other South Asians.We report that the Ror are the modern population that is closest to the first prehistorical and early historical South Asian ancient samples near the Indus Valley, and they also harbor the highest Steppe-related, EHG, and Neolithic Anatolian ancestry. However, compared to other adjoining groups, the Ror show less affinity with the Neolithic Iranians. The Ror population can plausibly be used as an alternative proxy for ANI in future demographic modeling of South Asian populations.

The bar graph below explains it very well, where it can be seen that the proportion of the steppe orange component is higher among Rors and Jats than either the Pathans, the Brahmins or any other South Asian group.

The admixture proportions as per the qpAdm given in the Supplementary Table 11 and it is instructive to observe that the steppe_emba proportion for Rors is estimated at 57 % of total ancestry while for Jats it is 61 %. The same proportions for Brahmins from UP, Gujarat & Bengal are 46 %, 45 % & 44 % respectively. Even for Pashtuns from Afghanistan it is 52 % and for Kalash it is 58 %. Only the Yaghnobis and Pamiris from Central Asia are estimated to have a higher proportion of steppe_EMBA at 62 % & 67 % respectively.

Before moving forward it is necessary to point out that the light blue component observed in the admixture graph which is highest among the Northern Europeans is not the same as the steppe_EMBA or steppe_MLBA ancestry. Steppe_EMBA & Steppe_MLBA are an amalgation of the light blue, the dark blue (Anatolian-Farmer related) and the light green (Iran_N/CHG) components you see in the admixture graphs. So while the light blue component which peaks in Northern Europe is significantly less among South Asians, the light green component which correlates well with Iran Neolithic type ancestry, peaks in South Asia but it present at a lot less proportion among the northern Europeans.

Infact, the authors even stress that,

The Ror and Jat peoples stand out for having the highest proportion of Steppe_ MLBA ancestry (- 63%). The proportion of Steppe ancestry in the Ror is similar to that observed in present day Northern Europeans

Therefore, the predominance of the light blue component in Northern Europeans is not alone an indication that their ‘steppe’ ancestry is far higher than among South Asians.


Now, if steppe-related ancestry correlates with presence and spread of Indo-European languages, the above data clearly implies that the highest steppe-related and therefore IE ancestry among South Asians is among the Jats  & Rors, significantly higher than in other NW groups as well as Brahmins and Kshatriyas. Jats and Rors sampled for the study, live in Haryana & Western UP, which is the Vedic homeland.

It therefore supports the ancient Indian tradition according to which the region of Haryana & Western UP was the homeland of the Vedic people from where they spread out across Northern India. It can therefore be argued perfectly well, that the Brahmins and Kshatriyas in other regions have higher proportion of ‘steppe’ ancestry than the lower classes around them precisely because they have greater percentage of their ancestry derived from the ‘steppe’ rich people from the Vedic homeland. It has long been an argument that the ‘steppe’ ancestry in higher among the Brahmins and Kshatriyas than the lower castes across all regions of India  and that this was evidence of IE culture spreading in South Asia with the ‘steppe’ ancestry. But the example of Jats and Rors in Haryana puts to doubt all such claims. Instead, we can argue that the higher ‘steppe’ related ancestry in Upper Castes across India is a function of them having a greater portion of their ancestry from their Vedic forefathers who lived in Haryana & Western UP, just as is suggested by the Vedic tradition.


I may finally add that there is a closely related group based on close fst distances and similar admixture proportions that likely descends from the core group that was responsible for the spread of this ancestry into the Caucasus and the steppe. This group consists of Rors, Jats, Kalash, Pashtun, Pathan, Tajik & Pamiri. They have broadly similar levels of Iran_N (15 to 30 %), Steppe_EMBA (49 to 67 %) & Onge (15 to 25 %) as per the qpAdm modelling in table S11. Fst distances also indicate that they are quite closely related. For example, the Fst distance between Rors and Pamiris (0.0069), Pashtuns (0.0057) & Tajiks (0.0058) is similar to Fst distances of Rors with neighbouring groups like Kamboj (0.0088), Gujjar (0.0064), Khatri (0.0056), Brahmins (0.0052) & Kshatriyas (0.0062). Considering the fact that Rors (& perhaps Jats) haven’t probably admixed with Pamiris, Tajiks or Pashtuns since millenia, their Fst distances would have been even less initially. The other Indus Valley modern populations are also not very far off in terms of Fst distances with each other but the above groups seem to form a subset among them.

https://i.imgur.com/TrNPI1r.jpg

It is conceivable that an ancestral group related to these populations with similar levels of ancestry proportions as exhibited by them (but perhaps with lowel levels of AASI – since BMAC has only 5 % in comparison to Pamiris who have 15 %), spread out from North India to Central Asia and those from Central Asia venturing further towards Caucasus and from there onto the steppe.

 

2+